1. Simpson, C. and J. B. C. Jackson, 2022. Bryozoan Revelations, Science Advances. 8 (13), eabp9344. dooi:10.1126/sciadv.abp9344

  2. Leslie, A.B., Simpson, C., and Mander, L. 2021. Pulsed rise in plant complexity linked to reproductive innovations. Science 373: 1368–1372.

  3. Simpson, C. 2021. Adaptation to a viscous Snowball Earth Ocean as a path to complex multicellularity. American Naturalist. 198:5.

  4. Laurel Sky Hiebert, Leandro M. Vieira, Stefano Tiozzo, Carl Simpson, Richard K. Grosberg, Alvaro E. Migotto, Andre C. Morandini, Federico D. Brown Simpson, C. (2021) From the individual to the colony: marine invertebrates as models to understand levels of biological organization. J Exp Zool (Mol Dev Evol).

  5. Simpson, C. 2020. An ecological driver for the macroevolution of morphological polymorphism within colonial invertebrates. J Exp Zool (Mol Dev Evol). DOI: 10.1002/(ISSN)1552-5015

  6. Hiebert, LS, Simpson, C, Tiozzo, S. 2020. Coloniality, clonality, and modularity in animals: The elephant in the room. J Exp Zool (Mol Dev Evol). 1– 14.

  7. Simpson, C., A Herrera-Cubilla, and J.B.C Jackson. 2020. How colonial animals evolve. Science Advances 6 : eaaw9530

  8. Simpson, C. 2018. Serving up light. Current Biology 28(16): R873-R875

  9. Hopkins, M. J., D. W. Bapst, C. Simpson, R. C. M. Warnock, 2018. The inseparability of sampling and time and its influence on attempts to unify the molecular and fossil records. Paleobiology, 44(4): 561-574. doi:10.1017/pab.2018.27

  10. Simpson, C. 2017. The measurement of species selection on evolving characters. bioRχiv.

  11. Simpson, C. 2017. The case for species selection. bioRχiv doi:

  12. Simpson, C., Jackson, J. B. C., Herrera-Cubilla, A. 2017. Evolutionary determinants of morphological polymorphism in colonial animals. American Naturalist 190(1):17-28. Preprint available at bioRχiv doi:

  13. Simpson, C. 2016. Understanding macroevolution through the origin of higher taxa. Ecology, 97: 3246–3248. doi:10.1002/ecy.1550

  14. Soreghan, G.S., Heavens, N.G., Hinnov,L.A., Aciego, S.M., and Simpson, C.. 2015. Reconstructing the dust cycle in deep time: The case of the late Paleozoic icehouse. In, Earth-Life Transitions: Paleobiology in the context of Earth System Evolution. P. David Polly, Jason J. Head, and David L. Fox, editors. The Paleontological Society Papers volume 21. pp. 83 - 120

  15. Orzechowski, E.A., R. Lockwood, J.E. Byrnes, S.C. Anderson, S. Finnegan, Z.V. Finkel, P.G. Harnik, D.R. Lindberg, L.H. Liow, H.K. Lotze, C.R. McClain, J.L. McGuire, A. O’Dea, J.M. Pandolfi, C. Simpson, D.P. Tittensor. Marine extinction risk shaped by trait-environment interactions over 500 million years. 2015. Global Change Biology. 21(10): 3595–3607. doi: 10.1111/gcb.12963

  16. Finnegan, S., Anderson, S.C., Harnik P.G., Simpson, C., Tittensor, D.P., Byrnes, J.E., Finkel, Z.V., Lindberg, D.R., Liow, L.H., Lockwood, R. Lotze, H.K., McClain, C.M., McGuire, J.L., O’Dea, A., Pandolfi, J.M. (2015) Paleontological baselines for evaluating extinction risk in the modern oceans. Science. 348(6234): 567-570. DOI: 10.1126/science.aaa6635

  17. Simpson, C. and Erwin, D.H., 2014. Enriching macroevolution. Science. 344(6189): 1234-1235.

  18. Hopkins, M., Simpson, C., and Kiessling, W., 2014. Differential niche dynamics among major marine invertebrate clades. Ecology Letters. 17(3):314-323. doi: 10.1111/ele.12232.

  19. Nowak, M. D., Smith, A. B., Simpson, C., and Zwickl, D. J. 2013. A Simple Method for Estimating Informative Node Age Priors for the Fossil Calibration of Molecular Divergence Time Analyses. PLoS ONE 8(6): e66245. doi:10.1371/journal.pone.0066245.

  20. Simpson, C. 2013. Species selection and the macroevolution of coral photosymbiosis and coloniality. Evolution 67(6): 1607-1621 doi://10.1111/evo.12083

  21. Kiessling, W., Simpson, C., Beck, B., Mewis, H. & Pandolfi, J. 2012. Equatorial decline of reef corals during the last Pleistocene interglacial. PNAS. 109(52): 21378-21383. doi:10.1073/pnas.1214037110

  22. Harnik, P. G., Simpson, C., & Payne, J. L. 2012. Long-term differences in extinction risk among the seven forms of rarity. Proceedings of the Royal Society B - Biological Sciences. doi:10.1098/rspb.2012.1902

  23. Harnik, P. G., Lotze, H., Anderson, S. C., Finkel, Z. V. Finnegan, S., Lindberg, D. L., Liow, L. H., Lockwood, R., McClain, C. M., McGuire, J. L., O’Dea, A., Pandolfi, J. M., Simpson, C., Tittensor, D. 2012. Extinctions in ancient and modern seas. Trends in Ecology and Evolution. 27(11) 608-617. doi://10.1016/j.tree.2012.07.010

  24. Simpson, C., Müller, J., 2012. Species selection in the molecular age, in From clone to bone: the synergy of morphological and molecular tools in paleobiology, Johannes Müller and Rob Asher, eds. Cambridge University Press. p. 116-134.

  25. Simpson, C. 2012. The evolutionary history of division of labour. Proceedings of the Royal Society B - Biological Sciences. 279(1726): 116-121. doi:10.1098/rspb.2011.0766.

  26. Simpson, C., Kiessling, W., Mewis, H., Baron-Szabo, R. C. & Müller, J. 2011. Evolutionary diversification of reef corals: a comparison of the molecular and fossil records. Evolution. 65(11) 3274–3284. DOI: 10.1111/j.1558-5646.2011.01365.x

  27. Liow, Lee Hsiang, Carl Simpson, Frédéric Bouchard, John Damuth, Benedikt Hallgrimsson, Gene Hunt, Daniel W. McShea, Jeffery Powell, Nils Christian Stenseth, Melissa Stoller & Gunter Wagner. 2011. Pioneering Paradigms and Magnificent Manifestos – Leigh Van Valen’s priceless contributions to evolutionary biology. Evolution. 65(4) 917-922. DOI: 10.1111/j.1558-5646.2011.01242.x

  28. Simpson, C. 2011. How many levels are there? How insights from evolutionary transitions in individuality help measure the hierarchical complexity of life. In, The Major Transitions in Evolution Revisited, Brett Calcott & Kim Sterelny eds. MIT Press.

  29. McShea, D. W. & Simpson, C. 2011. The Miscellaneous Transitions in Evolution. In, The Major Transitions Revisited, Brett Calcott & Kim Sterelny eds. MIT Press.

  30. Kiessling, W. & Simpson, C. 2011 On the potential for ocean acidification to be a general cause of ancient reef crises. Global Change Biology. 17(1): 56 - 67. doi: 10.1111/j.1365-2486.2010.02204.x

  31. Simpson, C. & Kiessling, W. 2010. The diversity of life through time. In: Encyclopedia of Life Sciences (ELS). John Wiley & Sons, Ltd: Chichester.

  32. Simpson, C. & Kiessling, W. 2010 The role of extinction in large-scale diversity-stability relationships. Proceedings of the Royal Society B - Biological Sciences 277(1686): 1451-1456.

  33. Simpson, C. 2010 Species selection and driven mechanisms jointly generate a large-scale morphological trend in monobathrid crinoids. Paleobiology 36(3): 481-496.

  34. Kiessling, W., Simpson, C. & Foote, M. 2010 Reefs as Cradles of Evolution and Sources of Biodiversity in the Phanerozoic. Science 327(5962): 196-198.

  35. Simpson, C. & Harnik, P. G. 2009 Assessing the role of abundance in marine bivalve extinction over the post-Paleozoic. Paleobiology 35(4): 631–647.

  36. Alroy, J., Aberhan, M., Bottjer, D., Foote, M., Fürsich, F. T., Harries, P. J., Hendy, A. J. W., Holland, S., Ivany, L. C., Kiessling, W., Kosnik, M. A., Marshall, C. R., McGowan, A. J., Miller, A., Olszewski, T. D., Patzkowsky, M., Peters, S. E., Villier, L., Wagner, P., Bonuso, N., Borkow, P. S., Brenneis, B., Clapham, M. E., Fall, L. M., Ferguson, C. A., Hanson, V. L., Krug, A. Z., Layou, K. M., Leckey, E. H., Nürnberg, S., Powers, C. M., Sessa, J. A., Simpson, C., Tomašových, A. & Visaggi, C. C. 2008 Phanerozoic trends in the global diversity of marine invertebrates. Science 321(5885), 97-100.

  37. Sánchez, J. A., Lasker, H. R., Zeng, W., Coluci, V. & Simpson, C. 2003 How similar are branching networks in nature? A view from the ocean: Caribbean gorgonian corals. Journal of Theoretical Biology 222, 135-138.